Phacopidae

Abstract: Phacopid trilobites are relatively widespread in Devonian deposits of north Gondwana and some have been collected from several sections of the Saoura Valley in SW Algeria. New occurrences of phacopids assigned to Trimerocephalus, Dianops and Phacops sensu lato are described from this area and comparisons are made with closely allied species from Morocco and Europe. The trilobite assemblages of the ‘Argiles de Marhouma’ Formation are considered to be Frasnian–Famennian in age. These new occurrences have been integrated into an analysis of Upper Devonian phacopid biodiversity. Diversity fluctuations reflect environmental changes, bioevents and stratigraphic turnovers throughout the Upper Devonian. Peak diversity was attained after the post-Kellwasser event.

Abstract: A nautiloid conch containing many disarticulated exoskeletons of Omegops cornelius (Phacopidae, Trilobita) was found in the Upper Devonian Hongguleleng Formation of the northwestern margin of the Junggar Basin, NW China. The similar number of cephala, thoraces and pygidia, unbroken thoraces, explicit exuviae, and lack of other macrofossils in the conch, indicate that at least seven individual trilobites had moulted within the nautiloid living chamber, using the vacant chamber of a dead nautiloid as a communal place for ecdysis. This exuvial strategy manifests cryptic behaviour of trilobites, which may have resulted from the adaptive evolution induced by powerful predation pressure, unstable marine environments, and competition pressure of organisms occupying the same ecological niche in the Devonian period. The unusual presence of several trilobites moulting within a nautiloid conch is possibly associated with social behaviours in face of a serious crisis. New materials in this study open a window for understanding the survival strategy of marine benthic organisms, especially predator-prey interactions and the behavioural ecology of trilobites in the middle Palaeozoic.