TL;DR: Preliminary phylogenetic and morphological evidence suggest that tooth-digging behavior arose at least twice in North American beavers.
Abstract: Migmacastor procumbodens, gen. et sp. nov. is described from the late Arikareean of Nebraska based on a nearly complete skull. The new taxon exhibits various incisal specializations (e.g., incisors...
TL;DR: A new tribe of Castoroidinae, Nothodipoidini, is proposed for Nothoides and Microdipoides Korth and Stout, based on the fossorial adaptations of the skull, and their generally smaller size.
Abstract: The skull and mandible of Nothodipoides Korth are described. There are a number of adaptations present that are shared with other rodents with tooth-digging behavior (procumbent, elongated, and flattened incisors; anteriorly tilted occipital; strongly arched upper diastema). A new tribe of Castoroidinae, Nothodipoidini, is proposed for Nothodipoides and Microdipoides Korth and Stout, based on the fossorial adaptations of the skull, and their generally smaller size. It appears that fossorial adaptations have occurred in castorids at least three times: the Palaeocastorinae (Whitneyan through Arikareean), the primitive beaver Migmacastor (Arikareean), and the Nothodipoidini (late Barstovian through the Clarendonian).