Abstract: Scandinavia was one of the last geographic areas in Europe to become habitable for humans after the Last Glacial Maximum (LGM). However, the routes and genetic composition of these postglacial migrants remain unclear. We sequenced the genomes, up to 57× coverage, of seven hunter-gatherers excavated across Scandinavia and dated from 9,500–6,000 years before present (BP). Surprisingly, among the Scandinavian Mesolithic individuals, the genetic data display an east–west genetic gradient that opposes the pattern seen in other parts of Mesolithic Europe. Our results suggest two different early postglacial migrations into Scandinavia: initially from the south, and later, from the northeast. The latter followed the ice-free Norwegian north Atlantic coast, along which novel and advanced pressure-blade stone-tool techniques may have spread. These two groups met and mixed in Scandinavia, creating a genetically diverse population, which shows patterns of genetic adaptation to high latitude environments. These potential adaptations include high frequencies of low pigmentation variants and a gene region associated with physical performance, which shows strong continuity into modern-day northern Europeans.

Abstract: The extent to which prehistoric migrations of farmers influenced the genetic pool of western North Africans remains unclear. Archaeological evidence suggests that the Neolithization process may have happened through the adoption of innovations by local Epipaleolithic communities or by demic diffusion from the Eastern Mediterranean shores or Iberia. Here, we present an analysis of individuals’ genome sequences from Early and Late Neolithic sites in Morocco and from Early Neolithic individuals from southern Iberia. We show that Early Neolithic Moroccans (∼5,000 BCE) are similar to Later Stone Age individuals from the same region and possess an endemic element retained in present-day Maghrebi populations, confirming a long-term genetic continuity in the region. This scenario is consistent with Early Neolithic traditions in North Africa deriving from Epipaleolithic communities that adopted certain agricultural techniques from neighboring populations. Among Eurasian ancient populations, Early Neolithic Moroccans are distantly related to Levantine Natufian hunter-gatherers (∼9,000 BCE) and Pre-Pottery Neolithic farmers (∼6,500 BCE). Late Neolithic (∼3,000 BCE) Moroccans, in contrast, share an Iberian component, supporting theories of trans-Gibraltar gene flow and indicating that Neolithization of North Africa involved both the movement of ideas and people. Lastly, the southern Iberian Early Neolithic samples share the same genetic composition as the Cardial Mediterranean Neolithic culture that reached Iberia ∼5,500 BCE. The cultural and genetic similarities between Iberian and North African Neolithic traditions further reinforce the model of an Iberian migration into the Maghreb.

Abstract: In this contribution we dismantle the perceived role of marine resources and plant foods in the subsistence economy of Holocene foragers of the Central Mediterranean using a combination of dental calculus and stable isotope analyses. The discovery of fish scales and flesh fragments, starch granules and other plant and animal micro-debris in the dental calculus of a Mesolithic forager dated to the end of the 8th millenium BC and buried in the Vlakno Cave on Dugi Otok Island in the Croatian Archipelago demonstrates that marine resources were regularly consumed by the individual together with a variety of plant foods. Since previous stable isotope data in the Eastern Adriatic and the Mediterranean region emphasises that terrestrial-based resources contributed mainly to Mesolithic diets in the Mediterranean Basin, our results provide an alternative view of the dietary habits of Mesolithic foragers in the Mediterranean region based on a combination of novel methodologies and data.

Abstract: Near Eastern Neolithic farmers introduced several species of domestic plants and animals as they dispersed into Europe. Dogs were the only domestic species present in both Europe and the Near East prior to the Neolithic. Here, we assessed whether early Near Eastern dogs possessed a unique mitochondrial lineage that differentiated them from Mesolithic European populations. We then analysed mitochondrial DNA sequences from 99 ancient European and Near Eastern dogs spanning the Upper Palaeolithic to the Bronze Age to assess if incoming farmers brought Near Eastern dogs with them, or instead primarily adopted indigenous European dogs after they arrived. Our results show that European pre-Neolithic dogs all possessed the mitochondrial haplogroup C, and that the Neolithic and Post-Neolithic dogs associated with farmers from Southeastern Europe mainly possessed haplogroup D. Thus, the appearance of haplogroup D most probably resulted from the dissemination of dogs from the Near East into Europe. In Western and Northern Europe, the turnover is incomplete and haplogroup C persists well into the Chalcolithic at least. These results suggest that dogs were an integral component of the Neolithic farming package and a mitochondrial lineage associated with the Near East was introduced into Europe alongside pigs, cows, sheep and goats. It got diluted into the native dog population when reaching the Western and Northern margins of Europe.

Abstract: The multi-period (~38,000–6000 cal BP) site of Franchthi Cave, located in the Argolid peninsula of southern mainland Greece, is unique in the Eastern Mediterranean for preserving a long archaeological sequence extending from the Upper Palaeolithic through to the end of the Neolithic period. In this paper, we present new anthracological (carbonized fuel wood waste) evidence from Franchthi Cave with which we reconstruct the changing ecology of woodland vegetation in its environs during the late Pleistocene and the early-mid Holocene. The integrated archaeobotanical record (charred wood and non-wood macro-remains) demonstrates that in the Lateglacial the now-submerged coastal shelf of the southern Argolid peninsula was covered by steppe grassland vegetation dominated by junipers, almonds, cereals and legumes. The rapid climatic amelioration that marked the start of the Holocene brought about the disappearance of juniper and the expansion of deciduous woodland, cereals and lentils. This woodland-grassland biome bears no analogues in the modern and historical vegetation ecology of the Aegean basin. Instead, it is directly comparable to the steppe woodland biomes exploited by late Pleistocene and early Holocene hunter-gatherers in Southwest Asia, and points to the convergent evolution of late Pleistocene and early Holocene plant exploitation strategies between the two regions. Continuous sea-level rise during the early Holocene led to the gradual extinction of this unique palaeohabitat, which acted as the catalyst for the selective introduction of domesticated cereal crops at Franchthi Cave in the early 9th millennium cal BP. Our meta-analysis of the non-wood archaeobotanical data puts into question the concept of the wholesale introduction of a crop “package” by pioneer settler groups arriving from the East. It is proposed instead that selective cereal crop introduction formed part of a complex pattern of sociocultural interactions that brought together indigenous and immigrant groups into new communities.

Abstract: In 1981 one of us (Cherry) first attempted to identify spatial and temporal patterning in the human colonization of the Mediterranean islands. Since the 1980s, slowly accumulating evidence has suggested that the Mediterranean islands were sporadically inhabited by hunter-gatherers during the Late Pleistocene and Early Holocene. Here we seek to establish whether or not these data exhibit regularity. We suggest that evidence for Upper Palaeolithic to Mesolithic activity, tending to cluster on larger or less remote islands, indicates that while humans were clearly capable of reaching the Mediterranean islands prior to the Neolithic, their general reluctance to do so can be explained in terms of the variable environmental attractiveness of the insular Mediterranean. Tending to be relatively small, dry, and biologically depauperate, the Mediterranean islands were largely inhospitable to mobile groups preferring extensive territories with diverse and robust biotas. Sedentism only became a widely viable ...

Abstract: The development of farming was a catalyst for the evolution of the human diet from the varied subsistence practices of hunter-gatherers to the more globalised food economy we depend upon today. Although there has been considerable research into the dietary changes associated with the initial spread of farming, less attention has been given to how dietary choices continued to develop during subsequent millennia. A paleogenomic time transect for 5 millennia of human occupation in the Great Hungarian Plain spanning from the advent of the Neolithic to the Iron Age, showed major genomic turnovers. Here we assess where these genetic turnovers are associated with corresponding dietary shifts, by examining the carbon and nitrogen stable isotope ratios of 52 individuals. Results provide evidence that early Neolithic individuals, which were genetically characterised as Mesolithic hunter-gatherers, relied on wild resources to a greater extent than those whose genomic attributes were of typical Neolithic European farmers. Other Neolithic individuals and those from the Copper Age to Bronze Age periods relied mostly on terrestrial C3 plant resources. We also report a carbon isotopic ratio typical of C4 plants, which may indicate millet consumption in the Late Bronze Age, despite suggestions of the crop’s earlier arrival in Europe during the Neolithic.

Abstract: The purpose of this thesis is to evaluate and deduce the varied lifeways of Early Holocene foragers in southern Scandinavia. By taking an interdisciplinary approach, zooarchaeological data have been applied to the study of different aspects of Early and Middle Mesolithic subsistence, in order to frame a discussion concerning our current understanding of culture and life in early north European societies. Three different sites/areas are the focus: Norje Sunnansund, Huseby Klev and Gotland/Gisslause. However, all available material from the chosen temporal and spatial frame have been incorporated to enable holistic discussions. The three focus areas combined comprise all available coastal settlements with well-preserved organic material from the Early Mesolithic period, which has led to discussions centred on the use of aquatic resources and the importance of fish.To address the different aspects of Early and Middle Mesolithic subsistence, multiple approaches have been taken, whereby zooarchaeological methods have been combined with statistical, chemical, physical and ethnographical tools for analysis. The focus has varied between fish storage and conservation practice, by presenting evidence for delayed-return subsistence strategies through means of large-scale fish fermentation, and discussions concerning the evidence for a delayed-return lifestyle and sedentism, through the study of zooarchaeological assemblages. Furthermore, taphonomy is highlighted and discussed in order to address the many biases affecting the recovery of freshwater fish bones and the consequences for detecting a freshwater fish-based diet. Pioneer subsistence strategies are studied, and changes through time are highlighted in marine coastal regions. In addition, the reservoir effect in radiocarbon dating (14C) of human bones has been examined to evaluate the consequences of a freshwater reservoir effect stemming from a large dietary input of freshwater fish. Furthermore, stable isotopes values, δ13C and δ15N, in the collagen from all available Early and Middle Mesolithic humans have also been analysed and modelled, in order to evaluate the importance of each individual protein source in the diet.The results from the different approaches taken indicate that humans relied on fish to a higher degree and from an earlier date than previously assumed. This has implications for how Early Holocene societies are interpreted; indicating the use of delayed-return subsistence strategies, diminishing mobility and emerging sedentism already existed during the Early Mesolithic period. Overall, the results of this thesis suggest a growing territoriality, which implies that the emergence of social stratification is conceivable at an early stage of Scandinavian prehistory and offers an insight into the lifestyle of Early Holocene foragers at latitudes around 55–59° N.

Abstract: Small game seems to have increased during the Upper Palaeolithic to the detriment of large game on the Iberian Peninsula. The economical and socio-cultural factors associated with this ecological s...

Abstract: The fauna of Neolithic Lameiras includes abundant sheep. Many could be secure- ly identified by applying criteria described by the late Joachim Boessneck as well as metrical methods. Sheep bones from Early Neolithic contexts, several dated directly via 14C, pinpoint the arrival here, 5450 cal BC, of this exotic animal three thousand years after its domestication 5000 km to the east. Thus sheep were transported at a rate of 1,6 km per year – considerably faster than suggested by the ‘wave of advance’ theory. It therefore seems probable that part of the journey was undertaken by ship. Most of the mammal remains identified at Lameiras belonged to domes- ticated forms and besides sheep and some goat, they include cattle and pig. Zooarchaeologically there is little difference between Early and Late Neolithic. However the Neolithic spectrum of species contrasts with that from a small assemblage in the underlying Mousterian level as well as other pre-Neolithic assemblages in Portugal. It is possible that in southern Portugal the adop- tion of animal husbandry was sudden. Measurements of the remains of Canis, Bos, Ovis, Capra and Sus compared with an increasingly large corpus of data from the South-Western part of the Iberian Peninsula indicate several occasions when these animals underwent size changes. Bos, Capra and Canis were considerably larger in the Pleistocene – a size difference now documented in other regions. Besides a Pleistocene-Holocene reduction in size, they underwent a further dim- inution associated with their domestication. It is possible that aurochs and wild boar recovered some of their former size after the Neolithic, perhaps due to a relaxation of hunting pressure after the Mesolithic. Domestic sheep, goats and cattle increased in size in more recent times perhaps reflecting Moslem and Christian improvements. KEYWORDS: FAUNA, DOMESTICATES, NEOLITHIC, PORTUGAL, EVOLUTION

Abstract: Burials of the Late Palaeolithic (14 000-11 600 cal years before present, henceforth BP) are a rare phenomenon in Europe. Several sites possess burials of single and double individuals. As with the preceding Magdalenian, the burial of more than two individuals in the same grave cutting seems to be unusual, but does occur occasionally. The deposition of isolated and disarticulated human remains with or without cut marks seems additionally to belong to the Magdalenian context. In the final Palaeolithic phase (13 000-11 600 cal years BP) there is evidence for cemetery-like clusters of burials, which contrast to the Magdalenian evidence, instead showing some similarities with the succeeding Mesolithic. The earliest Mesolithic burials 11 600-10 500 cal BP) are a very rare phenomenon, covering a short time span between the beginning of the Preboreal and the beginning of the Boreal phase of the early Holocene. Here the evidence includes single inhumations, cemetery-like structures and a number of isolated human remains. Caves and rock shelters were the most common places for inhumations in both the final Palaeolithic and the early Mesolithic. Although the number of sites with a chronological continuity from the LUP to the Early Mesolithic burial is low, several aspects indicate a general continuity in burial patterns over this period. Apart from this continuity, the Mesolithic burials in general seem to represent a new level of diversity in burial practices.This article is part of the theme issue 'Evolutionary thanatology: impacts of the dead on the living in humans and other animals'.

Abstract: During the Neolithic period, cattle were used not only for their meat and their milk but also for their strength. Unfortunately, apart from the discovery of specific instruments (yokes, travois, wheels, ards, etc.), it is not easy to demonstrate archaeologically their use for work. Nevertheless, the bone pathologies related to this activity can be analyzed. The methodological approach employed in this study is based on multivariate analyses (PCA) applied to the dimensions of the first and second phalanxes, as well as to shape indices of the same bones determined by the Mosimann method. The measurements of aurochs and domestic cattle bones from nineteen Mesolithic and Neolithic sites form the data matrix. The results of this study attest, on the one hand, that cattle were used for draught and transport during the Neolithic in various parts of Europe and the Near East and, on the other hand, that this use and its corollary, castration, are practices that can be dated back earlier than is generally accepted

Abstract: This contribution reviews the evidence for technological continuity and change (both gradual and abrupt) among the classic Magdalenian, Azilian, Asturian and non-coastal Mesolithic culture-adaptive traditions against the backdrop of environmental change across the Pleistocene–Holocene (i.e. Bolling–Boreal) transition in Cantabrian Spain. It explores the interacting, non-exclusive roles of environment, demography, subsistence and ‘tradition’ or historical contingency in explaining both inertia and change in the cultural records (including settlement, mobility, artistic activity and even world-view or ideology) observed by archeologists in this long-occupied, southerly wing of the classic Franco-Cantabrian culture area.