TL;DR: Differences in the pattern of replacement of the preceding Galerix species in Germany and Spain suggest that ecological factors played a role in that replacement, while in Greece, G. symeondisi is not replaced by G. exilis.
Abstract: The distribution of the Early Miocene galericine Galerix symeonidisi, with records from Greece, Germany, Spain and Austria is discussed. The identification of G. cf. symeonidisi from the Anatolian Cankiri basin is changed into G. cf. uenayae on the basis of additional material. Galerix symeonidisi enters Europe at the MN 3/ MN 4 transition. Differences in the pattern of replacement of the preceding Galerix species in Germany and Spain suggest that ecological factors played a role in that replacement. Galerix symeonidisi is succeeded by G. exilis in Western and Central Europe. A G. symeonidisi - G. exilis lineage, as has been proposed in literature, is rejected. The record from the Daroca-Calamocha area (Spain) is instead interpreted as a case of gradual replacement. The replacement is considered the result of an opening of the landscape when conditions became dryer. In Greece, G. symeondisi is not replaced by G. exilis. In Karidia, a galericine has been found which shows features intermediate between G. symeondisi and the oldest representatives of Schizogalerix, suggesting that symeonidisi might be ancestral to that genus.
TL;DR: A new early species of Galerix is document in sites from Sehwan, the Zinda Pir Dome, the Potwar Plateau, and Banda Daud Shah ranging in age from about 19 to 14 Ma, previously based on a few teeth from Daud Khel.
Abstract: Hedgehogs (erinaceid insectivores) are a common element in Miocene small mammal faunas of Pakistan, but little material has been formally described. Here, we report on extensive collections from numerous localities across Pakistan, most from the Potwar Plateau, Punjab, and the Sehwan area in Sindh. The dominant erinaceid is Galerix, which is also known from Europe, Turkey, and East Africa. We document a new early species of Galerix, Galerix wesselsae, in sites from Sehwan, the Zinda Pir Dome, the Potwar Plateau, and Banda Daud Shah ranging in age from about 19 to 14 Ma. This species briefly co-occurs with and is then replaced by the smaller Galerix rutlandae, previously based on a few teeth from Daud Khel. G. rutlandae is a common element of Chinji Formation faunas through about 11.5 Ma. Scanty material of an apparently new species of Schizogalerix succeeds Galerix. After 10.2 Ma, no galericines occur in Pakistan, but rare erinaceines are later faunal components.
TL;DR: Comparison of the late early to early late Miocene record of the Iberian insectivores shows that, particularly during the late Early Miocene, central European taxa have transient occurrences in Spain, while most taxa appear earlier and survive longer in central Europe.