TL;DR: Study of new material of this fossil species confirms that it is a stem-group salmonine, with a mixture of primitive and derived salmonine features in its skull, but with its postcranial skeleton essentially of modern salmonine construction.
TL;DR: Seven genera—Brachymystax, Acantholingua, Salmothymus, Hucho, Salvelinus, Salmo, and Oncorhynchus—make up the living Salmoninae, and monophyly of recognized genera is consistent with all 12 estimates.
Abstract: Seven genera—Brachymystax, Acantholingua, Salmothymus, Hucho, Salvelinus, Salmo, and Oncorhynchus—make up the living Salmoninae. Relationships of 33 extant and 4 fossil salmonid species and subspecies were studied on the basis of 119 characters analyzed by parsimony algorithms. Twelve equally parsimonious trees each requiring 253 steps were calculated. Monophyly of recognized genera is consistent with all 12 estimates. The earliest branch of the family Salmonidae is the subfamily Coregoninae. Its sister group is the clade including the Thymallinae and Salmoninae. Within the Salmoninae, Eosalmo, from the Eocene of British Columbia, is the sister group of all living genera, as previously shown by Mark Wilson. The living Asian species Brachymystax lenok is the sister species of all other living Salmoninae, as documented by Carroll Norden. Three species of archaic trouts from the Mediterranean area—Acantholingua ohridana, Salmothymus obtusirostris, and Salmothymus (Platysalmo) platycephalus—branch of...