TL;DR: The skull of a small anomodont therapsid, from the Tapinocephalus Assemblage Zone in Northern Cape Province, South Africa, represents a new basal dicynodont, and a phylogenetic analysis indicates that Colobodectes is the basalmost member of a dicYNodont clade that excludes Eodicynodon.
Abstract: The skull of a small anomodont therapsid, from the Tapinocephalus Assemblage Zone (Abrahamskraal Formation, Beaufort Group, Upper Permian) in Northern Cape Province, South Africa, represents a new basal dicynodont and is described in detail Colobodectes cluveri gen et sp nov is distinguished from other dicynodonts by an anteroposteriorly extensive caniniform process, parietals that were broadly overlapped posterolaterally by posterodorsal processes of the postorbitals, diverging anterior palatal ridges, and a dorsoventrally low foramen magnum A phylogenetic analysis indicates that Colobodectes is the basalmost member of a dicynodont clade that excludes Eodicynodon This position is not particularly strong, as two additional steps are needed to make Colobodectes and Eodicynodon oosthuizeni exchange places on the most parsimonious tree Another discovery of the phylogenetic analysis is that there is little basis for recognizing Eodicynodon oelofseni as the closest relative of E oosthuizeni The former species is identified as the sister taxon of a clade that includes the latter and all other dicynodonts
TL;DR: In this article, a comprehensive analysis of the fauna indicates that three vertebrate assemblage zones existed in the Upper Permian of China and that the Dashankou lower tetrapod fauna consisting mainly of therapsids is the most primitive and abundant one.
Abstract: A comprehensive analysis of the fauna indicates that three vertebrate assemblage zones existed in the Upper Permian of China. Among them the Dashankou lower tetrapod fauna consisting mainly of therapsids is the most primitive and abundant one. It is more closely related to Zone II of Russia than to the Eodicynodon Assemblage Zone of South Africa. Zone II of Russia in the Late Permian played a role of bridge for the dispersal of tetrapods from North America to South Africa and China at that time.
TL;DR: The Beaufort Group of South Africa has been known for their tetrapod fossil record, which is the richest and most complete Middle Permian to Middle Triassic record for any terrestrial sequence in the world as discussed by the authors.
Abstract: The fluvio-lacustrine rocks of the Beaufort Group, South Africa have long been known for their tetrapod fossil record, which is the richest and most complete Middle Permian to Middle Triassic record for any terrestrial sequence in the world. The abundance of fossil material has enabled the Beaufort Group to be biostratigraphically subdivided into between 8 and 10 tetrapod assemblage zones, of which the lowest three (Eodicynodon, Tapinocephalus and Pristerognathus) are attributed to the Middle Permian. These lower assemblage zones record the earliest therapsiddominated faunas and, because they were recorded during a largely uninterrupted period of deposition, make the Beaufort Group the only place in the world where biodiversity change through the terrestrial Middle Permian can be effectively studied.
TL;DR: A new specimen of Eodicynodon oosthuizeni, discovered near Prince Albert Road in the Western Cape province of South Africa, has a double distinction: the presence of two canines on the left maxilla is considered to be pathological and not the result of tooth replacement as discussed by the authors.
Abstract: A new specimen of Eodicynodon oosthuizeni, discovered near Prince Albert Road in the Western Cape province of South Africa, has a double distinction. Of stratigraphic importance is the fact that it is the only specimen of Eodicynodon known from above the first maroon mudrocks of the Beaufort Group, a feature which has important implications for understanding the development of the earliest terrestrial environments of Gondwana during the middle Permian. Of anatomical significance is the presence of two canines on the left maxilla. This rare condition, known in only three dicynodonts, is here considered to be pathological and not the result of tooth replacement.