TL;DR: The new specimen of T. riograndensis from the Carnian of the Candelária Sequence (Southern Brazil) is described and represents one of the largest specimens of a carnivorous probainognathian from the Late Triassic known to date and contributes to knowledge of size variation in ecteniniids.
Abstract: Ecteniniidae comprises an endemic radiation of carnivore probainognathian cynodonts from the Late Triassic of South America. Three taxa have been included in this clade: Ecteninion lunensis Martinez et al., 1996 and Diegocanis elegans Martinez et al., 2013 from Argentina, and Trucidocynodon riograndensis Oliveira et al., 2010 from Brazil. Herein, a new specimen (skull and mandible) assigned to T. riograndensis from the Carnian of the Candelaria Sequence (Southern Brazil) is described. A phylogenetic analysis recovered the new specimen as the sister taxon of the holotype of T. riograndensis, and both in a trichotomy with E. lunensis and D. elegans, all supporting the monophyly of Ecteniniidae. The new specimen of T. riograndensis is almost 20% larger than its holotype. Therefore, it represents one of the largest specimens of a carnivorous probainognathian from the Late Triassic known to date and contributes to knowledge of size variation in ecteniniids.
TL;DR: The nearly complete skull of a recently discovered carnivorous eucynodont, Ecteninion lunensis, gen. et sp.
Abstract: The nearly complete skull of a recently discovered carnivorous eucynodont, Ecteninion lunensis, gen. et sp. nov., is described. This is the first new genus of carnivorous cynodont reported from the Late Triassic (middle Carnian) Ischigualasto Formation of northwestern Argentina, and is unique to the formation. Diagnostic features include partially differentiated cheek teeth that overlap in an imbricate pattern, elongate braincase and parietal crest, pronounced lateral expansion of the braincase, sagittal crest that overhangs the occipital plate in dorsal view, gracile postorbital bars, short osseous secondary palate, maxilla excluded from the border of the subtemporal fenestra, palatine and orbitosphenoid that contribute to the medial orbital wall, post-temporal foramen enclosed within the tabular, squamosals that nearly meet at the posterior end of the sagittal crest, narrow and deep groove for the external auditory meatus, jaw articulation located at the anterior base of the zygomatic arches, a...
TL;DR: An almost complete skeleton of a new carnivorous cynodont from the Upper Triassic of Southern Brazil, Trucidocynodon riograndensis gen. nov., is described and the overall morphology of the astragalocalcaneal complex is similar to that of Diademodon and of the therocephalian Bauria.
Abstract: An almost complete skeleton of a new carnivorous cynodont from the Upper Triassic of Southern Brazil, Trucidocynodon riograndensis gen. nov. et sp. nov., is described. The new taxon is very similar to Ecteninion lunensis Martinez et al. 1996 from Upper Triassic of Argentina (Ischigualasto Formation). Both have an elongated skull, large pterygoid flanges, a well developed orbitosphenoid, a relatively short osseous secondary palate, greatly developed canines and sectorial postcanines with posteriorly directed cusps. However, the new taxon shows some differences relative to E. lunensis such as an open pterygoparoccipital foramen, the posterior opening of the post-temporal foramen enclosed by tabular and squamosal, and upper incisors with serrated cutting edges. The more remarkable features of the postcranium of T. riograndensis are the presence of 32 presacral vertebrae, a greater number than in most of non-mammaliaform cynodonts, the accentuated lumbarisation of the posterior trunk vertebrae and a remarkable morphological gradient in the caudal vertebrae; the femur and humerus show some interesting adaptations suggesting a more upright limb posture. The overall morphology of the astragalocalcaneal complex is similar to that of Diademodon and of the therocephalian Bauria.