TL;DR: A cladistic analysis of 32 taxa (two gorgonopsians, seven therocephalians and 23 cynodonts) and 96 craniodental characters places P. elegans as sister taxon of Eucynodontia, and indicates paraphyly for Therocephalia, with the whaitsiid Theriognathus identified as sisterTaxon of CynodontIA.
Abstract: : Platycraniellus elegans is an enigmatic Triassic cynodont from South Africa that has only been briefly described previously. New preparation of the holotype and additional unpublished material allows a detailed redescription and comparison with different cynodonts. Platycraniellus elegans is recognized as a valid species of basal cynodont. The distinct suborbital angulation of the zygomatic process, previously considered as a diagnostic character in chiniquodontid cynodonts, and more recently observed in some galesaurids, is also present in P. elegans. A larger, second specimen was initially referred to P. elegans, but most recently considered to belong to Galesaurus planiceps. Close comparison of this specimen with the holotype of P. elegans and with galesaurid specimens allows a tentative allocation to G. planiceps. A cladistic analysis of 32 taxa (two gorgonopsians, seven therocephalians and 23 cynodonts) and 96 craniodental characters places P. elegans as sister taxon of Eucynodontia. Results from the analysis favour a dichotomy between (1) Cynognathia, including the sectorial-toothed cynodonts Ecteninion, Cynognathus and the gomphodont cynodonts, and (2) Probainognathia, including most sectorial-toothed eucynodonts (e.g. Lumkuia, Probainognathus, Chiniquodon), tritheledontids, tritylodontids and mammaliaforms. The Late Triassic sectorial-toothed Ecteninion is the most basal member of Cynognathia, whereas the Middle Triassic Lumkuia is the basal representative of Probainognathia. Tritylodontids (Oligokyphus and Kayentatherium) are placed among Probainognathia, forming a monophyletic group with the tritheledontid Pachygenelus, whereas Brasilitherium is the sister taxon of Mammaliaformes. The cladistic analysis also indicates paraphyly for Therocephalia, with the whaitsiid Theriognathus identified as sister taxon of Cynodontia.
TL;DR: A new trirachodontid cynodont is described from the base of the Burgersdorp Formation (Subzone A fauna of the Cynognathus Assemblage Zone), of the South African Karoo Basin and is characterized by circular to ovoid in outline upper postcanines and the absence of a maxillary platform lateral to the postcanine series.
TL;DR: The brain endocasts of the late Triassic traversodontids Siriusgnathus niemeyerorum and Exaeretodon riograndensis from southern Brazil are described based on virtual models generated using computed tomography scan data, showing an endocranial cavity that is not fully ossified.
Abstract: The brain endocasts of the late Triassic (Carnian) traversodontids (Eucynodontia: Gomphodontia) Siriusgnathus niemeyerorum and Exaeretodon riograndensis from southern Brazil are described based on virtual models generated using computed tomography scan data. Their skull anatomy resembles that of other non-mammaliaform cynodonts, showing an endocranial cavity that is not fully ossified. A "V-shaped" orbitosphenoid, neither fully developed nor ossified is present in E. riograndensis. The nasal cavity is confluent with the encephalic cavity. Thus, the anterior limit of the olfactory bulbs is not definite. The brain endocast is elongated, being narrow anteriorly and wide posteriorly, with the maximum width at the parafloccular cast. The olfactory bulbs do not present a clear division between their counterparts, due to the absence of a longitudinal sulcus. A longitudinal sulcus in the forebrain delimiting the cerebral hemispheres, the pineal tube, and the parietal foramen are absent in both taxa. The large and well-developed unossified zone is partially separated from the remaining endocast by a notch formed by the supraoccipital. The encephalization quotients, as well as the endocranial volume/body mass relationships of S. niemeyerorum and E. riograndensis are within the range expected for non-mammaliaform Therapsida.
TL;DR: A comparative analysis of the morphology of the nasal cavity, nasal recesses, nasolacrimal duct, and maxillary canals of both species using computed tomography highlights the changes that occurred in parallel to the origin of mammaliaforms.
Abstract: Traversodontidae is a group of Triassic herbivorous/omnivorous cynodonts that represents the most diversified lineage within Cynognathia. In southern Brazil, a rich fossil record of late Middle/mid-Late Triassic cynodonts has been documented, with Exaeretodon riograndensis Abdala, Barberena, and Dornelles, 2002 and Siriusgnathus niemeyerorumPavanatto et al., 2018 representing two abundant and well-documented traversodontids. The present study provides a comparative analysis of the morphology of the nasal cavity, nasal recesses, nasolacrimal duct, and maxillary canals of both species using computed tomography, highlighting the changes that occurred in parallel to the origin of mammaliaforms. Our results show that there were no ossified turbinals or a cribriform plate delimiting the posterior end of the nasal cavity, suggesting these structures were probably cartilaginous as in nonmammaliaform cynodonts. Both species show lateral ridges on the internal surface of the roof of the nasal cavity, but the median ridge for the attachment of a nasal septum is absent. Exaeretodon riograndensis and S. niemeyerorum show recesses on the dorsal region of the nasal cavity, which increase the volume of the nasal cavity, potentially enhancing the olfactory chamber and contributing to the sense of smell. On the lateral sides of the nasal cavity, the analyzed taxa show a well-developed maxillary recess. Although E. riograndensis and S. niemeyerorum have roughly similar nasal cavities, in the former taxon, the space between the left and right dorsal recesses of the nasal cavity is uniform along its entire extension, whereas this space narrows posteriorly in S. niemeyerorum. Finally, the nasolacrimal duct of S. niemeyerorum is more inclined anteroposteriorly than in E. riograndensis.
TL;DR: Cynognathus crateronotus is a species of large carnivorous cynodont, first named and best known from the Triassic Burgersdorp Formation (Beaufort Group, Karoo Basin) of South Africa.
Abstract: Cynognathus crateronotus is a species of large carnivorous cynodont, first named and best known from the Triassic Burgersdorp Formation (Beaufort Group, Karoo Basin) of South Africa. Fossils of the genus have also been reported from the upper Fremouw Formation of Antarctica, the Upper Omingonde Formation of Namibia, and the Rio Seco de la Quebrada Formation of Argentina. Without associated cranial material, however, distinguishing the postcrania of Cynognathus from that of the closely related cynognathian, Diademodon tetragonus, has proven difficult. Here we provide a more comprehensive diagnosis for Cynognathus crateronotus and describe two novel occurrences. First, parts of a medium-sized individual were recovered from a scrambled mass of dental and semiarticulated postcranial material from the lower Ntawere Formation of Zambia, a horizon that previously produced fossils of Diademodon and Kannemeyeria. Second, a large individual was collected from a recently discovered locality within the lower part of the Lifua Member of the Manda Beds of Tanzania. A large proportion of the postcranial skeleton, along with some craniodental remains, was found ex situ in a sandy streambed just downstream from a quarry that has produced the dicynodont Dolichuranus, an azendohsaurid archosauromorph, and the avemetatarsalian Teleocrater rhadinus. The widespread occurrence of Cynognathus across southern Pangea demonstrates its utility as a biostratigraphic marker, but recently published radiometric age estimates from Argentina suggest that either the genus persisted for over 10 million years or African strata traditionally interpreted as Middle Triassic are better understood as Late Triassic in age.