TL;DR: An upper Oligocene–lowermost Miocene Asiatic differentiation is suggested for the Elasmotheriine group, strengthened by a first appearance datum in the Dera Bugti area (lower part of the MN 3).
Abstract: The stratigraphical position of the Dera Bugti area (Baluchistan, Pakistan), which yielded the remains described herein, is discussed Dental and postcranial material attributed to Bugtirhinus praecursor gen et sp nov, from the lower Miocene (MN 3) of the same area, is described This species is the oldest and most primitive member of the elasmotheriine group within Rhinocerotidae (Mammalia) A new diagnosis is established for the Elasmotheriini Comparison with the other lower and middle Miocene elasmotheriine taxa leads to the tree [Bugtirhinus praecursor [Caementodon oettingenae [other elasmotheriine taxa]]] An upper Oligocene–lowermost Miocene Asiatic differentiation is suggested for the group, strengthened by a first appearance datum in the Dera Bugti area (lower part of the MN 3) The origin and dispersal of the elasmotheriines throughout Eurasia are discussed and correlated with major faunal exchanges Keywords: Bugti Hills, Pakistan, Rhinocerotidae, Elasmotheriini, Miocene, biostratigraphy, palaeobiogeography
TL;DR: A new outcrop, Hang Mon 2, provides evidence of the rhinoceroses Pleuroceros blanfordi and Bugtirhinus sp. as mentioned in this paper, and these fossils allow a correlation to the earliest Miocene (most probably ranging from ~23 to ~21 Ma; Aquitanian) based on faunal comparison with the Sulaiman Province of Pakistan.
Abstract: Current scientific knowledge of Tertiary fossils from south of the Ailao Shan-Red River shear zone is extremely poor, in sharp contrast with the situation nowadays, as the area of Laos and Vietnam is regarded as a global hotspot of biodiversity. In this context, the few localities that yielded fossil assemblages are of first importance for the understanding of Cenozoic palaeobiogeography and the tectonic and palaeogeographical evolution of the region. Hang Mon 1 (Son La Province, Northern Vietnam) was the first site that provided evidence of Tertiary mammals, but its age remained very controversial, interpretations ranging from Oligocene to Late Miocene. Herein, we re-investigate the mammal fauna of the locality based on newly collected material and previously published fossil mammals. A new outcrop, Hang Mon 2, provides evidence of the rhinoceroses Pleuroceros blanfordi and Bugtirhinus sp. Together with the earlier finds of uncommonly small-sized Protaceratherium, these fossils allow a correlation to the earliest Miocene (most probably ranging from ~23 to ~21 Ma; Aquitanian) based on faunal comparison with the Sulaiman Province of Pakistan. The revision of the mammals from Hang Mon 1 is in agreement with this stratigraphic proposal. In addition, the discoveries from Vietnam (the rhinocerotid assemblage and Hyotherium) further support the hypothesis of strong biogeographical and environmental affinities between Europe, the Indian Subcontinent and Southeast Asia (Vietnam) during the Aquitanian.