TL;DR: The relations of the giraffoids are assessed by methods of phylogenetic systematics and it is suggested that 'Palaeotragus' expectans and 'Palaelophrys' decipiens are closely related to Samotherium.
Abstract: Specimens of Climacoceras africanus are described from Maboko, Kenya. The new species Climacoceras gentryi is established on the basis of ossicones, mandibles, and upper and lower dentitions from Fort Ternan and Baringo, Kenya. By interpretation of its lower canines Climacoceras is identified as a giraffoid and is placed in the new family Climacoceridae. Canthumeryx sirtensis is identified from Muruarot and Rusinga, Kenya. A dentition and associated partial skeleton of this species are described. The teeth agree closely with specimens of the same species from Gebel Zelten, Libya. Zarafa zelteni from Gebel Zelten is synonymized with Canthumeryx sirtensis. Again on the basis of its lower canines Canthumeryx is identified as a giraffoid and is placed in the new family Canthumerycidae. Specimens of Palaeotragus primaevus are described from Baringo, Kenya. This material includes a cranium with the ossicones, skull roof, occipital and basicranial regions preserved. Palaeotragus primaevus specimens from Fort Ternan are used in this description and some of these are redescribed. The relations of the giraffoids are assessed by methods of phylogenetic systematics. Palaeomeryx, Prolibytherium and Propalaeoryx are excluded from the Giraffoidea as their lower canines are not known. The Palaeotraginae is shown to be an invalid polyphyletic grouping and the genus Palaeotragus is also shown to be polyphyletic. Palaeotragus microdon is probably synonymous with Palaeotragus rouenii and the three species Palaeotragus rouenii (P. microdon), Palaeotragus coelophrys and Palaeotragus quadricornis are retained in the genus Palaeotragus. It is suggested that 'Palaeotragus' expectans and 'Palaeotragus' decipiens are closely related to Samotherium. Palaeotragus primaevus is probably synonymous with Palaeotragus tungurensis and this species is closely related to the giraffines. With slight changes the subfamilies Sivatheriinae and Giraffinae are valid monophyletic groups. Hydaspitherium is synonymized with Bramatherium and the Sivatheriinae includes the genera Giraffokeryx, Birgerbohlinia, Bramatherium and Sivatherium while the Giraffinae includes the genera Honanotherium, Bohlinia and Giraffa and the species 'Palaeotragus' tungurensis (P. primaevus). Okapia is identified as the sister-group of the other giraffids. Triceromeryx is the sister-group of the Giraffidae. Canthumeryx is the sister-group of Triceromeryx plus the Giraffidae while Climacoceras is the sister-group of the other giraffoids.
TL;DR: It is concluded that the proximate ancestors of modern giraffes probably evolved in southern central Europe about 8 million years ago and may have arisen from the gelocid ancestral assemblage of 20–25 Mya via the family Palaeomerycidae.
Abstract: The origin, phylogeny, and evolution of modem giraffes (Giraffa camelopardalis) is obscure. We review here the literature and conclude that the proximate ancestors of modern giraffes probably evolved in southern central Europe about 8 million years ago (Mya). These ancestors appear to have arisen from the gelocid ancestral assemblage of 20–25 Mya via the family Palaeomerycidae. From the palaeomerycids arose the Antilocaprinae (Pronghorns) via the subfamily Dromomerycinae, and two subfamilies of giraffids, the Climacoceratidae and Canthumerycidae. The terminal genus of the Climacoceratid line was the now extinct massive giraffid Sivatherium sp. The Canthumerycids gave rise to the okapi and giraffes via the intermediate forms of Giraffokeryx, Palaeotragus sp. (of which the okapi is the extant form), Samotherium sp. and Bohlinia sp. All of which are extinct. Stimulated by climate change, progeny of Bohlinia entered China and north India, evolved into typical Giraffa species and became extinct there about 4 M...
TL;DR: Hadjidimovo and Kalimantsi in south-western Bulgaria yielded a rich collection of giraffes including Helladotherium duvernoyi, Bohlinia attica and Palaeotragus rouenii.
Abstract: The various late Miocene localities around the villages of Hadjidimovo and Kalimantsi in south-western Bulgaria yielded a rich collection of giraffes, including Helladotherium duvernoyi, Bohlinia attica and Palaeotragus rouenii. These are the same species as at Pikermi, suggesting that there is no great age difference between these localities, but the limb-bones of both former species are stouter at Kalimantsi than in the Greek site, a difference which is perhaps of ecological origin. The absence of Samotherium, known in the geographically close sites of Greek Macedonia might be a consequence of the isolation of the river basins.
TL;DR: The first results on the fossil mammalian fauna recovered during the first excava- tion season at the new site Pikermi Valley-1 (PV1) suggest a Turolian age.
Abstract: We present the first results on the fossil mammalian fauna recovered during the first excava- tion season at the new site Pikermi Valley-1 (PV1). The fauna comprises two hipparionine species (C. cf. mediterraneum, H. cf. brachypus), a giraffid (Bohlinia cf. attica), five bovids (Palaeoreas linder- mayeri, Protragelaphus skouzesi, Tragoportax cf. amalthea, Gazella sp., Bovidae indet.) and two carni- vores (? Adcrocuta eximia, Felidae indet.). The composition of the fauna suggests a Turolian age.
TL;DR: In this paper, Giraffidae specimens were discovered from the Late Miocene fossiliferous sector of Toros-Menalla (northern Chad) and assigned to Bohlinia, a genus known so far only in some late Miocene localities of Eurasia.