TL;DR: In this paper, the authors describe the development of the ovarian development in the black Beetle and the Heteronychus arator (Coleoptera : Scarabaeidae).
Abstract: (1976). Ovarian Development in Black Beetle, Heteronychus arator (Coleoptera : Scarabaeidae) New Zealand Entomologist: Vol. 6, No. 2, pp. 165-170.