TL;DR: The phylogenetic position of the derived Gondwanan theropod clade Unenlagiidae is analyzed, retrieves a monophyletic Avialae formed by UnenLagiidae plus Aves, and produces significant changes in the topological distribution within Deinonychosauria.
Abstract: In the present paper we analyze the phylogenetic position of the derived Gondwanan theropod clade Unenlagiidae. Although this group has been frequently considered as deeply nested within Deinonychosauria and Dromaeosauridae, most of the features supporting this interpretation are conflictive, at least. Modification of integrative databases, such as that recently published by Hu et al. (2009), produces significant changes in the topological distribution of taxa within Deinonychosauria, depicting unenlagiids outside this clade. Our analysis retrieves, in contrast, a monophyletic Avialae formed by Unenlagiidae plus Aves.
TL;DR: It is demonstrated that many features formerly regarded as being diagnostic of Avialae, including long and robust forelimbs, actually characterize the more inclusive group Paraves (composed of the avialans and the deinonychosaurs), which challenges the centrality of Archaeopteryx in the transition to birds.
Abstract: Archaeopteryx is widely accepted as being the most basal bird, and accordingly it is regarded as central to understanding avialan origins; however, recent discoveries of derived maniraptorans have weakened the avialan status of Archaeopteryx. Here we report a new Archaeopteryx-like theropod from China. This find further demonstrates that many features formerly regarded as being diagnostic of Avialae, including long and robust forelimbs, actually characterize the more inclusive group Paraves (composed of the avialans and the deinonychosaurs). Notably, adding the new taxon into a comprehensive phylogenetic analysis shifts Archaeopteryx to the Deinonychosauria. Despite only tentative statistical support, this result challenges the centrality of Archaeopteryx in the transition to birds. If this new phylogenetic hypothesis can be confirmed by further investigation, current assumptions regarding the avialan ancestral condition will need to be re-evaluated.