Arizonasaurus

Abstract: Postosuchus kirkpatricki (n.g., n.sp.) is a large, gracile, carnivorous thecodontian from the late Triassic Dockum Group of Texas. The species is characterized by its erect and facultative bipedal gait, highly advanced pelvis with slightly open acetabulum and footed pubis. The teeth are large, highly serrated, deeply set in sockets, and show a wave-like replacement pattern. Some flexibility in the palate and the lower jaw permitted the animal to swallow large chunks of flesh. Postosuchus was the arch predator of its time, and lived in the interfluves or upland regions along with aetosaurs, coelurosaurs, and ictidosaurs. The quarry's thanatocoenose probably reflects autochthonous burials of a Dockum community, perhaps in consequence of a catastrophic event such as a flash flood. Although Postosuchus exhibits many carnosaurian traits, its ankle joint is of \`crocodile-normal' pattern, which suggests its pseudosuchian affinity. All Triassic reptiles previously thought to be carnosaurs are considered herein as advanced thecodontians; true carnosaurs probably appeared in post-Triassic time. Postosuchus is grouped with Poposaurus, Teratosaurus, Arizonasaurus and the unnamed \`Warwick archosaur' in the family Poposauridae. The Poposauridae and Rauisuchidae are placed in the infraorder Rauisuchia. Tyrannosaurs probably evolved from poposaurids, as indicated by the correlation between skull, pelvis and ankle structure.

Abstract: Arizonasaurus babbitti is rediagnosed from a well-preserved partial skeleton from the early Middle Triassic Moenkopi Formation of northern Arizona. The skull, excluding the braincase, is described and compared to other “rauisuchians” and basal archosaurs. Two possible autapomorphies include a deep fossa hidden from lateral and medial views on the posteroventral edge of the ascending process of the maxilla and a tongue (pubic peduncle of the ilium) and groove (proximal portion of the pubis) articulation between the pubis and the ilium. The postcrania is described in detail. Tall neural spines form a sail that is similar to those of other basal archosaurs with unknown affinities (Ctenosauriscus, Lotosaurus, Bromsgroveia, “Hypselorhachis”). Additional characters in the pelvic girdle in Arizonasaurus unite this taxon with Ctenosauriscus, Lotosaurus, Bromsgroveia, and Hypselorhachus to form a group, the ctenosauriscids. Pelvic characters of the ctenosauriscids are also found in Poposaurus and “Chatterjeea”, su...

Abstract: The Moenkopi Formation has yielded partial and isolated remains of important archosaurs including rauisuchian skull fragments and isolated poposaur centra and pelvic girdle elements. A recently discovered skeleton referable to Arizonasaurus babbitti shows that most of these archosaurian remains belong to one taxon. Characteristics of the skeleton of Arizonasaurus show that it belongs to a poorly known group of Middle Triassic (240-230 Myr ago) archosaurs called the ctenosauriscids, and that ctenosauriscids are or are closely related to poposaurs. Furthermore, many characteristics of Arizonasaurus provide evidence that poposaurids and ctenosauriscids are derived rauisuchians. The presence of a poposaurid in the early Middle Triassic suggests that the divergence of birds and crocodiles occurred earlier than previously thought. Middle Triassic ctenosauriscids also allow the correlation of Triassic faunas in Europe, Asia, North America and Africa. The Moenkopi Formation fauna represents a transitional fauna between Early and Late Triassic faunas.

Abstract: The Lifua Member of the Manda Beds of Tanzania (Middle Triassic: ?latest Anisian) has yielded an exceptionally important assemblage of early archosaurs, including numerous well-preserved specimens. However, the majority of this material has not been described formally, frustrating attempts to incorporate it into studies on early archosaur diversity and evolution. We describe an anterior dorsal vertebra from the Lifua Member as the holotype of a new taxon, Hypselorhachis mirabilis. Hypselorhachis is characterised by the possession of an elongate neural spine that is at least 5.5 times the height of the centrum, and can be diagnosed on the basis of a single autapomorphy relating to the morphology of the prezygapophysis. Hypselorhachis is similar to other early Middle Triassic archosaurs with elongate neural spines, including Arizonasaurus, Ctenosauriscus, and Lotosaurus. It is possible that these taxa form a clade, Ctenosauriscidae, but further anatomical and phylogenetic work is required before th...