Turn your draft into a Biological Theory-ready manuscript: a 250-word abstract, 4–6 keywords and an anonymised main file for blind review, handled for you.
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Walter Veit
Marlize Lombard
Andrew Buskell
Elisabeth A. Lloyd
Authored by Sumalatha Gangadhar | Published on 28 September, 2026
Use this Biological Theory template when the manuscript has to match that journal before you upload it. Check the current rules in the Biological Theory author instructions; the Biological Theory journal page has the scope and latest issues.
It is the writing shell for Biological Theory: title and authors, the abstract, the body, declarations, figures, and the reference list. The limits that decide a desk reject are on the journal page linked above.
A Biological Theory submission has a title page with authors and affiliations, an abstract of up to 250 words, keywords, the main text for your article type, declarations, the reference list, and figures and tables.
Read the full rules on the journal page linked above before you upload.
The sections a Biological Theory submission is built from, in order: title page, abstract, keywords, main text, declarations, references, figures, and tables. It does not replace the journal's own checklist.
| Element | Biological Theory requirement |
|---|---|
| Abstract | An unstructured abstract of no more than 250 words |
| Main text | Original articles up to 7,000 words; Original papers up to 3,000 words; Original article of unusual significance up to 7,000 words |
| Declarations | Ethics approval/consent as applicable; competing interests; funding; data availability |
| Figures/tables | Publication-ready; cited in order |
| Supplemental | Upload separately when used |
| Keywords | 4–6 keywords |
| References | Citation style not stated in the accessible author guidelines; match recent articles and any editor instructions |
| Peer review | Double-anonymous (double-blind) peer review |
| Submission | www.editorialmanager.com/bthe |
These rules come from the Biological Theory author instructions. Journals change limits without notice, so check the current guide before you upload.
An unstructured abstract of no more than 250 words.
4–6 keywords.
Original articles up to 7,000 words; Original papers up to 3,000 words; Original article of unusual significance up to 7,000 words.
Double-anonymous (double-blind) peer review.
Reference managers format Biological Theory citations with the Springer - Basic (author-date) reference style, with author and year in the text and an alphabetical reference list. Zotero and Mendeley list it as "Biological Theory" in the open Citation Style Language database.
In-text citation: (Smith and Patel 2021) for a two-author article, (Garcia 2019) for a single-author book.
Reference list entries:
These examples show the pattern, not real sources. If the author guide gives a different rule for DOIs or for how many authors to list before et al., follow the guide.
Open the author guide once before you draft and once before you upload. Match a recent Biological Theory paper for depth, headings, and figure style. Check the word limit for your article type before you write the discussion, not after.
Then submit on the journal's portal, not from this page.