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The Company of Biologists Publication Guide

The Company of Biologists is a UK not-for-profit publishing charity founded in 1925 and based in Cambridge. It publishes five well-regarded biology journals - Development, Journal of Cell Science, Journal of Experimental Biology, Disease Models & Mechanisms, and Biology Open - and channels publishing surpluses back into the community through grants, meetings, and travelling fellowships.

Journals published: 5 - Development, JCS, JEB, DMM, Biology Open
Open access options: 2 fully-OA titles (DMM, Biology Open) + hybrid OA on the other 3
APC range: ~$2,500 - $5,400 per article (2026 fees; check current price pages)
Headquarters: Cambridge, United Kingdom
Read & Publish: Yes - uncapped fee-free OA for authors at participating institutions
Charitable mission: Surpluses fund grants, meetings, workshops, and travelling fellowships
Indexing: All five journals indexed in Scopus, Web of Science, and PubMed
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Publication Guide

The Company of Biologists is a not-for-profit publishing charity founded in Cambridge, UK, in 1925. Unlike commercial publishers, it exists to support biologists: surpluses from its five journals fund travelling fellowships, scientific meetings, workshops, and community sites such as the Node, preLights, and FocalPlane. Its portfolio is small but distinguished - Development, Journal of Cell Science (JCS), Journal of Experimental Biology (JEB), Disease Models & Mechanisms (DMM), and Biology Open (BiO) are each leading venues in their fields.

This guide covers how to choose among the five journals, the open-access routes and current APCs, the peer-review workflow, and how to submit.

Which Company of Biologists journal fits your paper?

You're most likely to...Choose this journal
Report advances in developmental biology or stem cellsDevelopment - the field's flagship since 1953 (originally the Journal of Embryology and Experimental Morphology); hybrid with an OA option.
Report advances in cell biologyJournal of Cell Science - a leading cell-biology journal with roots going back to 1853; hybrid with an OA option.
Report comparative animal physiology, biomechanics, or experimental zoologyJournal of Experimental Biology - the leading journal in comparative physiology, publishing since 1923; hybrid with an OA option.
Present a disease model or translational mechanism studyDisease Models & Mechanisms - fully Open Access, focused on model systems for human disease.
Publish methodologically sound biology without novelty-based selectionBiology Open - fully Open Access, sound-science criteria, the lowest APC in the portfolio.

What The Company of Biologists publishes

Five peer-reviewed journals, all in the biological sciences:

  • Development - developmental biology and stem-cell research; the group's highest-impact title.
  • Journal of Cell Science - all areas of cell biology, including a strong tools-and-resources section.
  • Journal of Experimental Biology - comparative animal physiology and biomechanics.
  • Disease Models & Mechanisms - fully-OA journal on model organisms and mechanisms of human disease, launched 2008.
  • Biology Open - fully-OA, launched 2011, publishing rigorous research across the biological sciences using soundness-based peer review.

All five are indexed in Scopus, Web of Science, and PubMed.

Open access and APCs

The Company of Biologists offers every major OA route:

  • Gold OA (fully-OA journals) - Disease Models & Mechanisms and Biology Open are 100% Open Access under CC BY.
  • Hybrid OA - Development, JCS and JEB offer an immediate-OA option per article; the publisher continues to run them as transformative journals, growing OA share year on year (cOAlition S ended its formal TJ framework at the end of 2024).
  • Subscription route (no fee) - articles in Development, JCS and JEB can be published at no cost and become free to read 12 months after publication; Green self-archiving of the accepted manuscript is permitted.
  • Read & Publish - institutional agreements give corresponding authors at hundreds of participating institutions uncapped, fee-free OA publishing across all five journals; an EIFL partnership extends fee-free OA to authors in 30+ developing and transition-economy countries.
JournalOA modelAPC (2026, USD, before tax)
DevelopmentHybrid~$5,400
Journal of Cell ScienceHybrid~$5,400
Journal of Experimental BiologyHybrid~$5,400
Disease Models & MechanismsFully OA~$4,250
Biology OpenFully OA~$2,500

Automatic full waivers apply for corresponding authors in low-income and lower-middle-income economies, and hardship waivers are available on request. Fees are revised periodically - always check the journal's open-access page before submitting.

Peer review process

  • Model: single-anonymized peer review across all five journals; Biology Open assesses soundness rather than perceived significance.
  • Editors: academic editors handle papers, supported by in-house professional editors.
  • Preprints: all five journals are preprint-friendly and integrate with bioRxiv, including direct transfer of manuscripts and reviews.
  • Transfers: rejected manuscripts can be transferred between Company of Biologists journals with reviews carried over, saving review cycles.

How to submit

  1. Pick the journal. Match your paper's field and selectivity expectations using the table above.
  2. Read the author guidelines. Each journal publishes detailed manuscript-preparation guidance at journals.biologists.com.
  3. Prepare your manuscript. Formatting is flexible at first submission; use the journal's SciSpace template to save formatting time. References follow the house numbered/author-date biology style per journal.
  4. Submit via Editorial Manager. The Company of Biologists migrated from its legacy BenchPress system to Editorial Manager in 2024; all new submissions now go through Editorial Manager, with bioRxiv transfer integration.
  5. Track and respond. Decisions, reviews, and revisions are handled in Editorial Manager.

Ethics and integrity

The Company of Biologists is a member of COPE, OASPA, and STM, and follows COPE guidelines on corrections and retractions. Data-availability statements are required, image-integrity screening is applied, and the charity's governance - a board of directors who are practising scientists - keeps editorial policy aligned with the research community.

Beyond the journals

A distinctive reason many biologists choose the Company: money stays in science. The charity awards travelling fellowships for lab visits, funds scientific meetings and workshops (including its own Journal Meetings and Workshops programme), and runs the community sites the Node (developmental biology), preLights (preprint highlights), and FocalPlane (microscopy).

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PUBLISHER COMPARISON

How The Company of Biologists compares

How The Company of Biologists compares across breadth of titles, open-access routes, APCs, waivers and turnaround.

The Company of Biologists
The Company of Biologists logo
How The Company of Biologists compares across breadth of titles, open-access routes, APCs, waivers and turnaround.
Rockefeller University Press
Rockefeller University Press logo
How Rockefeller University Press compares across breadth of titles, open-access routes, fees, waivers and turnaround.
EMBO Press
EMBO Press logo
How EMBO Press compares on breadth of titles, open-access routes, APCs, waivers and turnaround.
Core Features
Best for
Developmental biology, cell biology, comparative physiology, disease models
High-impact immunology, cell biology, and physiology research
High-profile molecular and cell biology, systems biology, molecular medicine
Typical APC (OA)
~$2,500 - $5,400
$2,000 - $6,000 (immediate OA); JHI $3,500; LSA $3,975
$8,270 flat across the four EMBO journals (2026)
LMIC fee waivers
Yes - automatic waivers for low/lower-middle-income economies + EIFL agreement
Yes - free Read-and-Publish policy for LMICs; JHI APCs auto-waived
Yes - discretionary waivers, country-tiered pricing pilot, EMBO YIP/GIN support
Read & Publish deals coverage
Strong for its size - hundreds of institutions worldwide, uncapped OA publishing
Germany, Ireland, Japan, Spain, UK, and US (Virginia) consortia
Via Springer Nature's institutional OA agreements
Free / subscription (non-OA option)
Yes - Development, JCS and JEB have a no-fee subscription route (free to read after 12 months)
Yes - JCB/JEM $2,500 flat fee; JGP no fee; free to read after 12 months
No - fully OA since 2024; Reviews and commissioned content carry no APC
Breadth of titles
5 journals, all in the biological sciences
5 journals, all life sciences - deliberately small and selective
5 journals, all life sciences - deliberately small and selective
Cheapest realistic OA route
Biology Open at ~$2,500 (or fee-free via Read & Publish)
JGP immediate OA at $2,000; JGP green route is fee-free
Life Science Alliance (co-published; lower APC - check its current price list)
Speed to first decision
Varies by journal; editor-led triage typically within days, reviews in weeks
Presubmission inquiries answered ~2 days; decisions by active scientists
Editorial pre-screen in days; peer-reviewed first decision typically 4-6 weeks
KEY BRANDS

The Company of Biologists Journal Family

All five Company of Biologists journals, each linked to its SciSpace journal page with metrics, templates, and submission details.

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Faqs

The Company of Biologists FAQs

The Company of Biologists is a UK not-for-profit publishing charity founded in 1925, based in Cambridge. It publishes five biology journals - Development, Journal of Cell Science, Journal of Experimental Biology, Disease Models & Mechanisms, and Biology Open - and reinvests publishing surpluses into grants, meetings, travelling fellowships, and community sites for biologists.