The authors describe carbon system formulation and simulation characteristics of two new global coupled carbon–climate Earth System Models (ESM), ESM2M and ESM2G. These models demonstrate good climate fidelity as described in part I of this study while incorporating explicit and consistent carbon dynamics. The two models differ almost exclusively in the physical ocean component; ESM2M uses the Modular Ocean Model version 4.1 with vertical pressure layers, whereas ESM2G uses generalized ocean layer dynamics with a bulk mixed layer and interior isopycnal layers. On land, both ESMs include a revised land model to simulate competitive vegetation distributions and functioning, including carbon cycling among vegetation, soil, and atmosphere. In the ocean, both models include new biogeochemical algorithms including phytoplankton functional group dynamics with flexible stoichiometry. Preindustrial simulations are spun up to give stable, realistic carbon cycle means and variability. Significant differences...

GFDL’s ESM2 Global Coupled Climate–Carbon Earth System Models. Part I: Physical Formulation and Baseline Simulation Characteristics

We examine the carbon balance of North American wetlands by reviewing and synthesizing the published literature and soil databases. North American wetlands contain about 220 Pg C, most of which is in peat. They are a small to moderate carbon sink of about 49 Tg C yr−1, although the uncertainty around this estimate is greater than 100%, with the largest unknown being the role of carbon sequestration by sedimentation in freshwater mineral-soil wetlands. We estimate that North American wetlands emit 9 Tg methane (CH4) yr−1; however, the uncertainty of this estimate is also greater than 100%. With the exception of estuarine wetlands, CH4 emissions from wetlands may largely offset any positive benefits of carbon sequestration in soils and plants in terms of climate forcing. Historically, the destruction of wetlands through land-use changes has had the largest effects on the carbon fluxes and consequent radiative forcing of North American wetlands. The primary effects have been a reduction in their ability to sequester carbon (a small to moderate increase in radiative forcing), oxidation of their soil carbon reserves upon drainage (a small increase in radiative forcing), and reduction in CH4 emissions (a small to large decrease in radiative forcing). It is uncertain how global changes will affect the carbon pools and fluxes of North American wetlands. We will not be able to predict accurately the role of wetlands as potential positive or negative feedbacks to anthropogenic global change without knowing the integrative effects of changes in temperature, precipitation, atmospheric carbon dioxide concentrations, and atmospheric deposition of nitrogen and sulfur on the carbon balance of North American wetlands.

/pdf/the-carbon-balance-of-north-american-wetlands-3dbcztv0vh.pdf

The carbon balance of north american wetlands

Subfossil pollen and plant macrofossil data derived from 14 C-dated sediment profiles can provide quantitative information on glacial and interglacial cli- mates. The data allow climate variables related to growing- season warmth, winter cold, and plant-available moisture to be reconstructed. Continental-scale reconstructions have been made for the mid-Holocene (MH, around 6 ka) and Last Glacial Maximum (LGM, around 21 ka), allowing comparison with palaeoclimate simulations currently being carried out as part of the fifth Assessment Report (AR5) of the Intergovernmental Panel on Climate Change. The synthesis of the available MH and LGM climate recon- structions and their uncertainties, obtained using modern- analogue, regression and model-inversion techniques, is presented for four temperature variables and two moisture variables. Reconstructions of the same variables based on

/pdf/pollen-based-continental-climate-reconstructions-at-6-and-21-q7dw2mep3u.pdf

Pollen-based continental climate reconstructions at 6 and 21 ka: A global synthesis

A working checklist of accepted taxa worldwide is vital in achieving the goal of developing an online flora of all known plants by 2020 as part of the Global Strategy for Plant Conservation. We here present the first-ever worldwide checklist for liverworts (Marchantiophyta) and hornworts (Anthocerotophyta) that includes 7486 species in 398 genera representing 92 families from the two phyla. The checklist has far reaching implications and applications, including providing a valuable tool for taxonomists and systematists, analyzing phytogeographic and diversity patterns, aiding in the assessment of floristic and taxonomic knowledge, and identifying geographical gaps in our understanding of the global liverwort and hornwort flora. The checklist is derived from a working data set centralizing nomenclature, taxonomy and geography on a global scale. Prior to this effort a lack of centralization has been a major impediment for the study and analysis of species richness, conservation and systematic research at both regional and global scales. The success of this checklist, initiated in 2008, has been underpinned by its community approach involving taxonomic specialists working towards a consensus on taxonomy, nomenclature and distribution.

/pdf/world-checklist-of-hornworts-and-liverworts-2eqzlt4g3g.pdf

World checklist of hornworts and liverworts.

. Peatlands contain a large belowground carbon (C) stock in the biosphere, and their dynamics have important implications for the global carbon cycle. However, there are still large uncertainties in C stock estimates and poor understanding of C dynamics across timescales. Here I review different approaches and associated uncertainties of C stock estimates in the literature, and on the basis of the literature review my best estimate of C stocks and uncertainty is 500 ± 100 (approximate range) gigatons of C (Gt C) in northern peatlands. The greatest source of uncertainty for all the approaches is the lack or insufficient representation of data, including depth, bulk density and carbon accumulation data, especially from the world's large peatlands. Several ways to improve estimates of peat carbon stocks are also discussed in this paper, including the estimates of C stocks by regions and further utilizations of widely available basal peat ages. Changes in peatland carbon stocks over time, estimated using Sphagnum (peat moss) spore data and down-core peat accumulation records, show different patterns during the Holocene, and I argue that spore-based approach underestimates the abundance of peatlands in their early histories. Considering long-term peat decomposition using peat accumulation data allows estimates of net carbon sequestration rates by peatlands, or net (ecosystem) carbon balance (NECB), which indicates more than half of peat carbon (> 270 Gt C) was sequestrated before 7000 yr ago during the Holocene. Contemporary carbon flux studies at 5 peatland sites show much larger NECB during the last decade (32 ± 7.8 (S.E.) g C m−2 yr–1) than during the last 7000 yr (∼ 11 g C m−2 yr–1), as modeled from peat records across northern peatlands. This discrepancy highlights the urgent need for carbon accumulation data and process understanding, especially at decadal and centennial timescales, that would bridge current knowledge gaps and facilitate comparisons of NECB across all timescales.

/pdf/northern-peatland-carbon-stocks-and-dynamics-a-review-3ngtefiod4.pdf

Northern peatland carbon stocks and dynamics: a review

along climatic, chemical, and physical gradients in peat lands of western Canada. -Nova Hedwigia 53: 27-7 1. Abstract: The bryophyte component of peat land veget ation is analysed for 395 stands on 335 study sites in western Canada. Sites are located along a broad transect from the Queen Charlotte Islands, British Columbia to southeastern Manitoba. The climate in the study area is divided into five zones : hyper­ oceanic, oceanic , suboceanic, subcont inental. A TWINSPAN anl\lysis separated the stands into six groups and selected indicator species for each group. A DCCA ordination showed that surf ace water che­

http://cfs.nrcan.gc.ca/bookstore_pdfs/11177.pdf

Bryophyte response surfaces along climatic, chemical, and physical gradients in peatlands of western Canada

The spatial development and vegetation history of a large boreal peatland complex in east-central Alberta was reconstructed to examine factors that control peatland development in continental regio...

https://cfs.nrcan.gc.ca/bookstore_pdfs/23819.pdf

Development of a peatland complex in boreal western Canada: lateral site expansion and local variability in vegetation succession and long-term peat accumulation

Fallen logs of trembling aspen (Populus tremuloides Michx.) and white spruce (Picea glauca (Moench) Voss) at various stages of decomposition were sampled from undisturbed and 1-, 14-, and 28-year-o...

Microfungus communities of white spruce and trembling aspen logs at different stages of decay in disturbed and undisturbed sites in the boreal mixedwood region of Alberta

Responses of northern peatlands to climate change: effects on bryophytes

The climate of North America and the adjacent ocean at 6000 BP was estimated using five independent approaches. Using pollen data, the terrestrial climate was estimated by the movement of ecozone boundaries and by the method of modern analogues. Both analyses indicate warmer temperatures in the western Great Lakes area and the northern Great Plains. A model of Sphagnum-dominated peatland initiation, when forced by Canadian Climate Model 6 ka output projected a cooler and (or) wetter climate for continental western North America. Contrary to this, a reconstruction of the distribution of Sphagnum-dominated peatlands in western Canada indicates that they were located north of their modern distribution, suggesting warmer and (or) drier conditions at 6000 BP. This interpretation is strengthened by observations of lower lake levels at 6000 BP in western Canada. This drier climate may have been associated with warmer conditions as indicated by the quantitative climate reconstructions. In general, eastern North A...

/pdf/the-climate-of-north-america-and-adjacent-ocean-waters-ca-6-25qu3zedx6.pdf

The climate of North America and adjacent ocean waters ca. 6 ka

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